Snowtrout
The common snowtrout or snowtrout (Schizothorax richardsonii), also known as "Asala", or "Asela" by the locals, is a species belong to the class of ray-finned fishes, family Cyprinidae, order Cypriniformes in the genus Schizothorax. The genus name Schizothorax is derived from Greek word, "schizein" meaning to divide, and "thorax" which means breast.
Common snowtrout is commonly found in the Himalayan region of India, Bhutan, Nepal, Pakistan, and Afghanistan. This species is adapted to fast-flowing, cold-water streams of Himalayan rivers. This species can be found in mountain rivers dwelling among rocks, feeding on algae, aquatic plants and detritus. They usually breed in April and May.
Common snowtrout has great commercial and cultural values to the local communities. They are much sought after as a consumption fish. However, their population is threatened by a number of factors including overfishing, pollution, the damming of rivers and the introduction of exotic fish, particularly salmonids and the population is declining.
Common snowtrout is the most common Schizothoracinae in the Himalayas. They can be found in almost all riverine systems of the Himalayas. They are distributed in India, Nepal, Bhutan, Pakistan, and Afghanistan. Water temperature is known to be the determining factor of geographical distribution and dispersal of common snowtrout. This species have the upper tolerance around 20 °C and are able to remain active in the near zero temperature in the streams of Lesser and Greater Himalayas during December and January.
Himalayan fish including common snowtrout spend most part of their lives swimming against the current. This behavior helps the, to maintain their upright position and to make respiration easier. The common snowtrout prefer to live in Himalayan streams and rivers especially rivers with moderate flow speed. They thrive in fast-flowing water by clutching onto the rocks and stones at the bottom of the streams. This species has the proper adaptation for the fast-flowing water such as the streamlined body which helps them move easily against the torrential flow of the streams. This streamlined body offer the least resistance to the fast current and enabling movement against the current more efficiently. The small scales reduce friction and the absence or reduction of lipidosis on the ventral part of their bodies enable to press themselves against the substrate. This species is herbivorous by feeding on algae and detritus from the rocky substrate. The paired fins (pectoral and pelvic) are large with thick and muscular bases. Pectoral and pelvic fins are close to each other to resist the current. The paired fins are horizontally positioned and are shifted to the sides so the ventral side can be attached to the rocks and act as adhesive organ. Schizothorax richardsonii have two pairs of barbels which are smaller and shorter relative to the eye diameter.The common snowtrout is reported to have territoriality behavior and has been observed to chase intruders to defend the limited food resource and available shelter. Such behavior develops after the young fish emerges from the eggs laid in gravel.
The breeding season for snowtrout has been recorded to occur between June and October with the peak in September. There have been observations indicating that Schizothoracines including S. richardsonii exhibit multiple spawning throughout the year following the temperature rise and flow rates which support egg laying.The female eggs are maturing (not fully mature) around February–May and ready to spawn around August–September, while the male sperms mature in March–April and ready to spawn in July–October. However, the fish spawn during spring despite the relatively immature gonads. The reason is the gonads fully mature in winter, but they are in dormant phase due to the low temperature, thus the fish spawn only on the returning favorable conditions in spring.The rise of temperature in streams from near freezing to 10-17 C during May – June induces common snowtrout to spawn. In Sutlej River, common snowtrout start the migration upstream along with the rise in water temperature in March. During the migration, the fish endure low water temperature of 8.0 – 9.5 °C due to the steady influx of snow-melt water. This low temperature induces the fish to migrate to side streams where the ground water temperature is around 17.5 – 21.5 °C to spawn. In other river, Ravi River, this species spawn in May. In the upper Beas, this species spawn only in July – August when the water temperature warms up to 16.5 – 18.5 °C. in the same drainage, common snowtrout migrate downstream to the lowermost reaches to spawn in October – December when the temperature is around 19 – 22.5 °C. Their eggs are large (3.0-4.0 mm diameter) and sticky. This species lay eggs in shallow pools of 50 –70 cm depth and remain adhered to the bottom until the fry hatch.
Threats to common snowtrout include habitat fragmentation, slow growth, poor disease resistance and low survival rates. It is believed that alien species introduction, such as the introduction of Brown Trout (Salmo trutta), is the threat to the persistence of this species. In response to climate change, it is predicted that there will be a range shift upward towards the higher elevation.
